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It's generally advisable to use the aircon at all times and best with it set on 'Auto', because by doing so you are likely to avoid unpleasant smells due to build up of bacteria etc. A further consideration is that the aircon coolant contains lubricants to keep the system working correctly, including lubrication of the system seals. If the seals dry the refrigerant can escape and you may be faced with an unwelcome bill to have them all replaced. I think you'll find a schedule for the air-conditioning service in your owners handbook - its usually at around two yearly intervals.
With current climate control systems the impact on performance and fuel consumption is much improved on what it would have been a few years ago.
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